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Vigilius (genus)
''Vigilius'' is an extinct genus of brachyopid temnospondyl amphibian from the Triassic of Arizona. It is known from the single type species ''Vigilius wellesi''.
The holotype specimen of ''Vigilius'' is a skull labelled UCMP 36199. The skull was found in an area alongside Arizona State Route 64 overlooking the Grand Canyon. It came from a layer of the Early Triassic Moenkopi Formation. The skull was described by paleontologists Samuel Paul Welles and Richard Estes in 1969 and attributed to a new genus and species of brachyopid called ''Hadrokkosaurus bradyi''. The holotype of ''Hadrokkosaurus'' was an isolated lower jaw. In 2000, paleontologists Anne Warren and Claudia Marsicano suggested that the lower jaw and skull represent two different species, as they were found over apart and come from two animals of different size. Warren and Marsicano assigned the skull the a new genus and species ''Vigilius wellesi''. The genus name comes from the Latin word ''vigilia'' meaning "keeping watch," a reference to its large eye sockets and "to the fact that the skull was keeping vigil over the Grand Canyon."〔 The species was named in honor of Welles. Several bones found around the Grand Canyon site were also attributed to ''Vigilius'', including a pterygoid bone that was described in 1971 as a scapulacoracoid. Additional remains were found west of Holbrook, Arizona.〔
==Phylogeny==
The material now assigned to ''Vigilius'' was first identified as remains of a brachyopoid. Warren and Marsicano conducted a phylogenetic analysis of many brachyopoids and found ''Vigilius'' to nest within the family Brachyopidae. Brachyopoidea is normally divided into the families Brachyopidae and Chigutisauridae, but these families are not always considered closely related; in some studies, Chigutisauridae is placed in a group of mostly Mesozoic temnospondyls called Stereospondyli while Brachyopidae is classified within a group of temnospondyls called Dvinosauria, which includes Carboniferous and Permian taxa that are older than ''Vigilius''. The relationship of dvinosaurians to other temnospondyls is uncertain, as they may or may not belong to Stereospondyli. Warren and Marsicano identified many features in ''Vigilius'' that are not seen in other brachyopoids but are common in dvinosaurians, including a narrow vomer bone at the front of the palate and a very wide cultriform process at the front of the parasphenoid, a bone on the underside of the skull. They considered ''Vigilius'' to be a dvinosaur in spite of the results of their phylogenetic analysis.〔
抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』
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